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Billard E, Serrão EA, Pearson GA, Engel CR, Destombe C, Valero M. Hybridisation in the Fucus spiralis /Fucus vesiculosus species complex (Fucaceae, Phaeophyceae): sexual phenotype and prezygotic fertility. European Journal of Phycology

Billard E, Daguin C, Pearson GA, Serrão, EA, Engel C, Valero M (2005) Genetic isolation between the three closely related taxa: Fucus vesiculosus, F.spiralis and F. ceranoides. Journal of Phycology 41: 900-905